In 2001, he moved his research program to the University of Missouri (MU) where he was the Gilbreath-McLorn Professor of Comparative Medicine, Director of the Comparative Medicine Center, Director of the Rat Resource and Research Center, and Chairman of the Veterinary Trichostatin A cell line Pathobiology Department. While at MU, he developed

three NIH-funded national animal resource centers which were focused, in large part, on comparative medicine and reproductive cryobiology. In collaboration with other faculty, John was instrumental in establishing the MU Mutant Mouse Resource and Research Center and the Rat Resource and Research Center both of which serve as critical repositories for valuable rodent models. John was also an active participant in establishing a similar resource for swine (National Swine Resource and Research Center). He was responsible for leadership and administration of the core groups involving novel clinical/translational methodologies, translational technologies/resources, and pilot and collaborative translational/clinical

studies. Most recently, Dr. Critser was awarded an R01 component of the Oncofertility U54 program, one of the first funded NIH Roadmap BMS-354825 nmr Initiative projects. Dr. Critser contributed greatly to our Society. He served as our Society President, member of Society Committees, on the Editorial Board of Cryobiology, and Chairman of the Society Annual Conference CRYO1997 and Co-Chair of CRYO2004. Dr. Critser was also a member of many other professional societies and editorial review boards; he was continuously funded by the NIH for over 20 years; and was the past chair of the NIH National Center for Research Resources (NCRR) Comparative Medicine Study Section. Dr. Critser was a well-respected scholar and researcher in the fields of cryobiology, comparative medicine and reproductive biology. He authored or Rucaparib co-authored over 190 publications. His vision and unique ability to forge fruitful and lasting collaborations among individuals with diverse expertise from all over the world were among his notable strengths.

More important to him than any of these other accomplishments, Dr. Critser was proud and passionate about training graduate students and post-doctoral fellows. He mentored more than 30 graduate students and 20 postdoctoral fellows, many of whom are now in professional and leadership roles in the areas of cryobiology, comparative medicine, reproductive biology, molecular biology, engineering, medicine and veterinary medicine. He not only nurtured them during their training but also continued to mentor, help, collaborate and support them as they matured professionally. John Critser was a devoted cryobiologist who contributed significantly to our field. While his career ended abruptly and far too soon, his contributions were reflective of someone with decades more time among us.